The story of two mice, one from the country and one from the city, who share their lifestyle with each other.

Triplets Jessica, JoAnn, and Jerzey and their friends in ballet class try to keep the building that houses The Nutcracker School from being torn down, while also trying to rescue a stray kitten.

Alexandrea is forced to leave her friends in Apple Creek when her mother insists on enrolling her at Harlem's Nutcracker School of Ballet, and while Al is already aware that she lacks dance skills, things get worse when she is chosen to be the Sugar Plum fairy in the school recital.

When Epatha tries to spice up the choreography of the new ballet in Harlem with her free-spirited style, the rest of the Sugar Plum ballerinas encourage her to keep her toes in line.