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Toula is a quiet, devoted daughter in a big, crazy Greek family. Working in her father's restaurant, she hides behind her family and keeps the world at a distance. One day at the restaurant she finds herself pouring coffee for a man who inspires her to change her life, and the way she sees the world ... forever.

After super villain Shredder escapes custody, he joins forces with mad scientist Baxter Stockman and two dimwitted henchmen, Bebop and Rocksteady, to unleash a diabolical plan to take over the world. As the Turtles prepare to take on Shredder and his new crew, they find themselves facing an even greater evil with similar intentions: the notorious Krang.